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Lepechinia hastata

(A.Gray) Epling (1940)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Asterids Order: Lamiales Family: Lamiaceae Genus: Lepechinia

pakaha, pitcher sage

Description

Key Characters:

Growth Form: Erect perennial herbs 8–15 dm tall, somewhat woody at base, glandular tomentose, the hairs branched.

Stems:

Roots:

Leaves: Leaves simple. Opposite. Blades hastate, 12–25(–35) cm long, 3.3–13 cm wide, leaves progressively smaller toward inflorescence, becoming ovate to lanceolate. Surfaces tomentose, but less so on upper surface; blade thick, chartaceous, minutely rugose. Margins crenulate. Petioles (lower leaves) 1.3–5.5 cm long; leaves nearer inflorescences sessile. Stipules absent.

Flowers: Flowers 5–15 in compound cymes, these arranged in open panicles, pedicels usually 8–13 mm long; each flower +/- bracteolate. Calyx slightly zygomorphic, 8–16 mm long, lobes subulate, closing at maturity, 5-toothed, the teeth subequal, sometimes coherent, splitting longitudinally, usually persistent or sometimes only ventral 1/2 persistent. Corolla zygomorphic, bilabiate, purplish red, slightly curved, 22–28 mm long, pubescent externally, tube funnelform, curved, exserted beyone calyx, puberulent within, 5-lobed, lobes broad and rounded, upper lip 2-lobed, lower lip 3-lobed, median lobe larger than lateral ones, emarginate, lateral lobes entire; nectary disk often present at base of ovary, annular or developed on anterior side only. Stamens 4, strongly exserted ca. 10–15 mm beyond corolla tube; anther sacs divergent; anthers dithecal, opening by longitudinal slits. Ovary superior, 2-carpellate; ovules 1 per ovary lobe, placentation basal-axile; style 1, exserted beyond corolla tube; stigmas 2.

Fruit: Nutlets 4; 1-seeded; black; ovoid; hard; the surface shiny to puberulent and smooth; ca. 4–5 mm long. Seeds with endosperm absent or scanty and oily.

Ploidy:

Habitat: Occurring in open shrubland or pastures.

Elevation Range: 610–1,220(–1,830) m.

Historical Distribution

Uses and Culture

USES

Natural History

Statewide Status

Indigenous or Naturalized

Island Status

O'ahu Only found in cultivation
Maui Indigenous or Naturalized

Dispersal Agents


Pollinators

Bibliography

Name Published In: Bull. Torrey Bot. Club 67: 511 (1940)

Other References

Wagner et al. 1990:804 (EM)

Occurrences

SNo. Scientific Name Scientific Name Authorship Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Lepechinia hastata (A.Gray) Epling Puu Nianiau, Haleakala PRESERVED_SPECIMEN Hosaka, E.Y. 1773 Maui BISH 1/28/1937
2 Lepechinia hastata (A.Gray) Epling omaopio, haleakala road grassland PRESERVED_SPECIMEN st.john, h. collector number: 24724 Maui BISH 12/24/1951
3 Lepechinia hastata (A.Gray) Epling Makawao, Haleakala Ranch rocky, open grass PRESERVED_SPECIMEN Hosaka, E.Y. 1680 Maui BISH 1/17/1937
4 Lepechinia hastata (A.Gray) Epling East Maui, Haleakala PRESERVED_SPECIMEN Mann, H. 401 Maui BISH 2/6/1905
5 Lepechinia hastata (A.Gray) Epling W Haleakala along road to summit; open scrub on lava PRESERVED_SPECIMEN Fosberg, F.R. 41525a Maui BISH 12/15/1961
6 Lepechinia hastata (A.Gray) Epling S Haleakala, below Puu Pane PRESERVED_SPECIMEN Forbes, C.N. 1843 M Maui BISH 3/4/1920
7 Lepechinia hastata (A.Gray) Epling East Maui, Haleakala open pasture PRESERVED_SPECIMEN Hjort, F. 10 Maui BISH
8 Lepechinia hastata (A.Gray) Epling Lyon Arboretum, middle lawn makai, front of cottage N PRESERVED_SPECIMEN Ishikawa, S. 68.308 Oahu BISH 7/20/1973
9 Lepechinia hastata (A.Gray) Epling Olinda, above Makawao with Ulex in pasture PRESERVED_SPECIMEN Fosberg, F.R. 55451 Maui BISH 4/15/1974
10 Lepechinia hastata (A.Gray) Epling Omaopio, Haleakala road grassland PRESERVED_SPECIMEN St. John, H. 24724 Maui BISH 12/24/1951
11 Lepechinia hastata (A.Gray) Epling east maui, makawao district, haleakala crater road PRESERVED_SPECIMEN thomas, p. collector number: s.n. Maui BISH 11/11/1995
12 Lepechinia hastata (A.Gray) Epling east maui, haleakala PRESERVED_SPECIMEN mann, h. brigham, w.t. collector number: 401 Maui BISH
13 Lepechinia hastata (A.Gray) Epling w haleakala along road to summit; open scrub on lava PRESERVED_SPECIMEN fosberg, f.r. collector number: 41525a Maui BISH 12/15/1961
14 Lepechinia hastata (A.Gray) Epling makawao, haleakala ranch rocky, open grass PRESERVED_SPECIMEN hosaka, e.y. collector number: 1680 Maui BISH 1/17/1937
15 Lepechinia hastata (A.Gray) Epling olinda, above makawao with Ulex in pasture PRESERVED_SPECIMEN fosberg, f.r. collector number: 55451 Maui BISH 4/15/1974
16 Lepechinia hastata (A.Gray) Epling east maui PRESERVED_SPECIMEN rock, j.f.c. collector number: 8553 Maui BISH 9/1/1910
17 Lepechinia hastata (A.Gray) Epling east maui, haleakala, near olinda Epiphytic on Halimeda discocdea PRESERVED_SPECIMEN forbes, c.n. collector number: 225 m Oahu BISH 7/1/1910
18 Lepechinia hastata (A.Gray) Epling east maui, haleakala pasture PRESERVED_SPECIMEN alexander, a.m. kellogg, l. collector number: 5293 Maui BISH 8/8/1947
19 Lepechinia hastata (A.Gray) Epling haleakala, 6000' PRESERVED_SPECIMEN Maui US 8/8/1967
20 Lepechinia hastata (A.Gray) Epling East Maui, S Haleakala, Olinda PRESERVED_SPECIMEN Forbes, C.N. 225 M Maui BISH 7/1/1910
21 Lepechinia hastata (A.Gray) Epling w slopes of haleakala along highway to summit Uncommon in dense Schinus thicket with Adiantum hispidulum. PRESERVED_SPECIMEN f. r. fosberg 41525a Lanai BISH 3/15/1961
22 Lepechinia hastata (A.Gray) Epling East Maui, Haleakala pasture PRESERVED_SPECIMEN Alexander, A.M. 5293 Maui BISH 4/1/1947
23 Lepechinia hastata (A.Gray) Epling East Maui, Haleakala Crater; W Kaupo Gap PRESERVED_SPECIMEN Medeiros, A.C. 423 Maui BISH 6/28/1983
24 Lepechinia hastata (A.Gray) Epling east maui, haleakala open pasture PRESERVED_SPECIMEN hjort, f. collector number: 10 Maui BISH
25 Lepechinia hastata (A.Gray) Epling east maui, s haleakala, olinda PRESERVED_SPECIMEN forbes, c.n. collector number: 225 m Maui BISH 7/1/1910
26 Lepechinia hastata (A.Gray) Epling lower slopes of haleakala PRESERVED_SPECIMEN h. mann & w. t. brigham 401 MU
27 Lepechinia hastata (A.Gray) Epling Olinda; collected near Olinda Endangered Species Facility, house of Fern Duvall. PRESERVED_SPECIMEN K. R. Wood 2869 Maui PTBG 11/15/1993
28 Lepechinia hastata (A.Gray) Epling E Maui, Olinda Homesteads above Makawao PRESERVED_SPECIMEN Fosberg, F. R. 55451 Maui US 4/15/1974
29 Lepechinia hastata (A.Gray) Epling haleakala PRESERVED_SPECIMEN wilkes explor. exped. s.n. Maui US 1838-08-08
30 Lepechinia hastata (A.Gray) Epling haleakala PRESERVED_SPECIMEN rock, j.f.c. Maui BISH 8/8/1918
31 Lepechinia hastata (A.Gray) Epling East Maui, Haleakala, near Olinda PRESERVED_SPECIMEN Forbes, C.N. 225 M Maui BISH 7/1/1910
32 Lepechinia hastata (A.Gray) Epling East Maui, Makawao District, Haleakala Crater Road PRESERVED_SPECIMEN Thomas, P. s.n. Maui BISH 11/11/1995
33 Lepechinia hastata (A.Gray) Epling east maui, haleakala crater; w kaupo gap PRESERVED_SPECIMEN medeiros, a.c. loope, l.l. collector number: 423 Maui BISH 6/28/1983
34 Lepechinia hastata (A.Gray) Epling lyon arboretum, middle lawn makai, front of cottage n PRESERVED_SPECIMEN ishikawa, s. collector number: 68.308 Oahu BISH 7/20/1973
35 Lepechinia hastata (A.Gray) Epling East Maui, Makawao District, between Pohakuokala and Waiale Gulches. In open pasturep plants apparently unbrowsed by domestic and feral animals. Vegetation: Dodonaea, Leptecophylla, Vaccinium, Sophora, Pteridium, Dryopteris, Asplenium, Coprosma, Carex, Morelotia, Sadleria, Luzula, Dubautia, Pellaea, Polystichum, Holcus, Ageratina, Pennisetum, Dactylis, Hypochoeris, Prunella, Lythrum. PRESERVED_SPECIMEN Oppenheimer, H.L. H20923 Maui BISH 2/11/2009
36 Lepechinia hastata (A.Gray) Epling Between Pohakuokala & Waiale gulches. In open pasture. PRESERVED_SPECIMEN Hank Oppenheimer H20923 Maui PTBG 2/11/2009
37 Lepechinia hastata (A.Gray) Epling sandwich islands, on mouna haleakala east maui PRESERVED_SPECIMEN wilkes expedition s.n. Hawaii US 1838-08-07
38 Lepechinia hastata (A.Gray) Epling honolulu, the lyon arboretum, middle lawn makai, in front of cottage h. PRESERVED_SPECIMEN s. ishikawa 269 Oahu US 7/20/1973
39 Lepechinia hastata (A.Gray) Epling PRESERVED_SPECIMEN w. b. hillebrand 346 Hawaii US
40 Lepechinia hastata (A.Gray) Epling Puu Nianiau, Haleakala PRESERVED_SPECIMEN Hosaka, E. Y. 1773 Maui US 1/28/1937
41 Lepechinia hastata (A.Gray) Epling e maui, olinda homesteads above makawao PRESERVED_SPECIMEN f. r. fosberg 55451 Maui US 4/15/1974
42 Lepechinia hastata (A.Gray) Epling PRESERVED_SPECIMEN Oahu US 11/19/1967
43 Lepechinia hastata (A.Gray) Epling hill mauka of olinda on way to summit of haleakala. PRESERVED_SPECIMEN o. degener 9512 Maui US 6/15/1927
44 Lepechinia hastata (A.Gray) Epling Honolulu, The Lyon Arboretum, in front of Cottage H., Middle Lawn Makai. PRESERVED_SPECIMEN Ishikawa, S. 269 Oahu US 7/20/1973
45 Lepechinia hastata (A.Gray) Epling Honolulu, The Lyon Arboretum, Middle Lawn Makai, In front of Cottage H. PRESERVED_SPECIMEN Ishikawa, S. 269 Oahu US 7/20/1973
46 Lepechinia hastata (A.Gray) Epling puu nianiau, haleakala PRESERVED_SPECIMEN e. y. hosaka 1773 Maui US 1/28/1937
47 Lepechinia hastata (A.Gray) Epling Haleakala Road, west slope. Dry brushland PRESERVED_SPECIMEN Fosberg, F. R. 9921 Maui US 8/23/1933
48 Lepechinia hastata (A.Gray) Epling puu nianiau, haleakala PRESERVED_SPECIMEN hosaka, e.y. collector number: 1773 Maui BISH 1/28/1937
49 Lepechinia hastata (A.Gray) Epling s haleakala, below puu pane PRESERVED_SPECIMEN forbes, c.n. collector number: 1843 m Maui BISH 3/4/1920
50 Lepechinia hastata (A.Gray) Epling East Maui PRESERVED_SPECIMEN Rock, J.F.C. 8553 Maui BISH 9/1/1910
51 Lepechinia hastata (A.Gray) Epling W slopes of Haleakala along highway to summit PRESERVED_SPECIMEN Fosberg, F. R. 41525a Maui US 3/15/1961
52 Lepechinia hastata (A.Gray) Epling probably maui PRESERVED_SPECIMEN j. f. rock 8553 Hawaii US
53 Lepechinia hastata (A.Gray) Epling honolulu, the lyon arboretum, in front of cottage h., middle lawn makai. PRESERVED_SPECIMEN s. ishikawa 269 Oahu US 7/20/1973
54 Lepechinia hastata (A.Gray) Epling s slope of haleakala, below puu pane PRESERVED_SPECIMEN c. n. forbes 1843 m. Oahu US 3/4/1920
55 Lepechinia hastata (A.Gray) Epling S slope of Haleakala, below Puu Pane PRESERVED_SPECIMEN Forbes, C. N. 1843 M. Maui US 3/4/2020
56 Lepechinia hastata (A.Gray) Epling Haleakala PRESERVED_SPECIMEN Wilkes Explor. Exped. s.n. Maui US 1838-00-00
57 Lepechinia hastata (A.Gray) Epling Hill mauka of Olinda on way to summit of Haleakala. PRESERVED_SPECIMEN Degener, O. 9512 Maui US 6/15/2027
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